Wednesday, January 3, 2018

"Researchers have identified around a dozen or so coyote vocalizations"........... "Some coyote sounds are used to defend their territory and dens and to tell other coyotes they’re around"............"There’s little evidence that vocalizations are used to coordinate pack hunting [or to celebrate the kill]"........... "Some research shows the alpha, or high-ranking, males and females and pairs do most of the vocalizing"..............."Howling sounds can travel around 1,000 yards and can be used by coyotes to identify who’s calling, their gender and perhaps their mood".......... Transient coyotes don’t usually vocalize as much as resident animals in order to avoid interactions"........... "Lone howls can also announce the location of an individual separated from their group"..........."The melodious cacophony and symphony of sounds(beau geste” effect) shouldn’t be used to claim that numerous coyotes are all over the place"............."Indeed, (most of us) overestimate the number of individuals actually present".........."We still have much to learn about coyote vocal communication"............ "Even after years of studying coyote calls, I was barely able to scratch the surface".......... "These tricksters hold their secrets tightly"

https://www.psychologytoday.com/blog/animal-emotions/201712/coyotes-dispelling-myths-about-who-they-are-what-they-do

CLICK LINK BELOW TO WATCH A LONE COYOTE HOWL AND YIP,,,,,,,,,,,IT OFTEN SOUNDS
LIKE SCORES OF COYOTES WHEN IT IS JUST ONE UTILIZING ITS AMAZING VOICE

https://youtu.be/YtsZoIe3Czk

Coyotes: Dispelling Myths About Who They Are, What They Do

12/21/17 Marc BeKoff  PH.D,

Coyote sounds have become increasingly more common in a wide range of urban landscapes, and while we know some details about why coyotes vocalize and what their different sounds mean, we really actually don’t know all that much.

What are coyotes talking about when they vocalize?
Recent claims about coyote behavior made by San Francisco Chronicle outdoor columnist Tom Stientsra prompted Camilla Fox, Founder & Executive Director of Project Coyote, and me, to write an op-ed for the San Francisco Chronicle titled "Coyotes howl to chat with their neighbors." Our op-ed, focusing on coyote vocalizations, reads as follows:
When you watch coyotes throw their heads back and sing to their heart’ content, they seem to enjoy it. That was Marc’s impression when he and his students studied wild coyotes in the Grand Teton National Park for more than eight years. It’s fun, it feels good, so why not howl?






But what are they saying?
Researchers have identified around a dozen or so coyote vocalizations. Some coyote sounds are used to defend their territory and dens and to tell other coyotes they’re around, but some vocalizations give away much more information.
There’s little evidence that vocalizations are used to coordinate pack hunting [or to celebrate the kill]. Some research shows the alpha, or high-ranking, males and females and pairs do most of the vocalizing.




Based on extensive and detailed research that involved recording and playing back howls and yips and observing the behavior of captive and free-ranging coyotes, wildlife researcher Philip Lehner 40 years ago placed coyote sounds into three general categories:
Greeting: Sounds include low-frequency whining, wow-oo-wowing (often called a greeting song), and group yip-howling (when reuniting and greeting).
Agonistic: These are vocalizations used during aggressive interactions and when coyotes display submission. They include woofing, growling, huffing (high-intensity threat), barking, bark-howling, yelping (submission and startle), and high-frequency whining (usually given by a subordinate coyote).




Contact: Sounds include lone howling (one of the most common vocalizations), group howling (when reuniting or in response to lone or group howls or yip howls), and group yip-howling (which may announce territory occupancy and may help regulate density of population).
Howling sounds can travel around 1,000 yards and can be used by coyotes to identify who’s calling, their gender and perhaps their mood. Transient coyotes don’t usually vocalize as much as resident animals in order to avoid interactions. Lone howls can also announce the location of an individual separated from their group
 One interesting and useful discovery is that humans aren’t very good at estimating how many coyotes are around by listening to their howls. Indeed, they overestimate the number of individuals actually present. So the melodious cacophony and symphony of sounds shouldn’t be used to claim that numerous coyotes are all over the place.
The more we understand all aspects of coyote behavior, the easier it will be to peacefully coexist with them. We should use what we know to protect them. State and local policies should embrace our understanding of coyote behavior.
We need facts, not myths
Sensationalist media often harms coyotes and other nonhuman animals. The fact remains that neither Camilla nor I could find any published studies that demonstrate "The louder the howl, the bigger the prize, often a fawn," or that "when a pack goes berserk in the middle of the night" it precedes hunting. Indeed, it would be nice to know more about coyote vocalizations in general. 
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 https://www.google.com/url?sa=t&rct=j&q=&esrc=s&source=web&cd=12&cad=rja&uact=8&ved=0ahUKEwi2iJ75rL3YAhVEU98KHeY7AicQFghdMAs&url=https%3A%2F%2Fnorthernwoodlands.org%2Foutside_story%2Farticle%2Fcoyotes&usg=AOvVaw0GjC-wEDu5zfAgdTntypmt

Coyotes: Listening to Tricksters



Coyotes: Listening to Tricksters
Illustration by Adelaide Tyrol
As the sunset colors fade from purple to black, the forest is dimly illuminated by a first quarter moon. An eerie sound breaks the calm. It is not the long, low, slow howling of wolves that can be heard further north, but the group yip-howl of coyotes: short howls that often rise and fall in pitch, punctuated with staccato yips, yaps, and barks.
When people hear coyote howls, they often mistakenly assume that they’re hearing a large pack of animals, all raising their voices at once.  But this is an auditory illusion called the “beau geste” effect. Because of the variety of sounds produced by each coyote, and the way sound is distorted as it passes through the environment, two of these tricksters can sound like seven or eight animals.
Group yip-howls are produced by a mated and territorial pair of “alpha” coyotes, with the male howling while the female intersperses her yips, barks, and short howls. “Beta” coyotes (the children of the alpha pair from previous years) and current year pups may join in if they are nearby, or respond with howls of their own.  And once one group of coyotes starts howling, chances are that any other alpha pairs nearby will respond in kind, with chorus after chorus of group yip-howls rippling across the miles.  
I spent seven years studying coyote vocal communication during my dissertation research at the University of California, Berkeley. While eastern coyotes are a larger and distinct subspecies from the western coyotes that I worked with, the basic findings of my research and the work done by others applies to all coyotes.  Coyotes have sometimes been called “song dogs,” and their long distance songs come in two basic types.
The first, the group yip-howl, is thought to have the dual purpose of promoting bonding within the family group while also serving as a territorial display. In other words, the coyotes are saying “we’re a happy family, and we own this turf so you better keep out.” In a sense, the group howls create an auditory fence around a territory, supplementing the physical scent marks left by the group.
Coyotes will also howl and bark separately. This second type of song is virtually always an indication of disturbance or agitation, and in my experience, the higher the proportion of howls, the more agitated the coyote is. Coyotes will howl and bark at neighbors who intrude on their territory, and at dogs, people, and other large animals that they perceive as a potential threat.
My research documented that coyote barks and howls are individually specific. Much like we can tell people apart by their voices, there is enough information in coyote vocalizations for me (OK, my computer if you want to get technical) to tell individuals apart. If, as I suspect, coyotes can distinguish each other by their song, it would not be analogous to the animals constantly shouting their own names; it would be more akin to our ability to recognize Marlon Brando because of the distinctive timbre and cadence of his voice. Characteristics including dominant pitch, duration, how quickly howls rise and fall in pitch, and tendency to “warble” while howling all distinguish one coyote from another.
For howls, this individual distinctiveness does not fade with distance. I was able to record and identify individual coyotes over a distance of greater than one mile. Given their keen hearing, it is likely coyotes can discern individual howls at much greater distances —three miles or more on a calm night.
Barks, on the other hand, degrade quickly over distance, with the higher frequencies fading first. This makes it theoretically possible for coyotes familiar with an individual (say, a mate or family group member) to determine roughly how far away that individual is, based on the proportion of high frequencies in the barks.
Imagine a scenario where a lone coyote is patrolling the territory boundary and comes across an intruder. He starts barking and howling, and his mate and beta children come running to the right place because his howls indicate how agitated he is, and his barks allow his family to pinpoint the direction and distance to his location. Although I was not able to prove that coyotes can do these tasks, the information needed is present in their calls and there are strong evolutionary advantages to learning how to use it.
We still have much to learn about coyote vocal communication. Even after years of studying coyote calls, I was barely able to scratch the surface. These tricksters hold their secrets tightly.

Brian Mitchell is an adjunct professor at the University of Vermont. Now that he has kids, that grad school schedule of getting up at 2 AM for field work sounds pretty relaxing.

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