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Coyotes-Wolves-Cougars.blogspot.com

Grizzly bears, black bears, wolves, coyotes, cougars/ mountain lions,bobcats, wolverines, lynx, foxes, fishers and martens are the suite of carnivores that originally inhabited North America after the Pleistocene extinctions. This site invites research, commentary, point/counterpoint on that suite of native animals (predator and prey) that inhabited The Americas circa 1500-at the initial point of European exploration and subsequent colonization. Landscape ecology, journal accounts of explorers and frontiersmen, genetic evaluations of museum animals, peer reviewed 20th and 21st century research on various aspects of our "Wild America" as well as subjective commentary from expert and layman alike. All of the above being revealed and discussed with the underlying goal of one day seeing our Continent rewilded.....Where big enough swaths of open space exist with connective corridors to other large forest, meadow, mountain, valley, prairie, desert and chaparral wildlands.....Thereby enabling all of our historic fauna, including man, to live in a sustainable and healthy environment. - Blogger Rick

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Thursday, September 25, 2014

It is so Important to remember that when regional Communities and Highway Traffic Departments finally decide to install widlife crosssings, they must build multiple "life path" structures at the agreed upon crossing point to provide connectivity for all species likely to use a given area(Little 2003)............ Different species prefer different types of structures (Clevenger et al. 2001; McDonald & St Clair 2004; Clevenger & Waltho 2005; Mata et al. 2005)................ For deer or other ungulates, an open structure such as a bridge is crucial................. For medium-sized mammals like black bears, and mountain lions, large box culverts with natural earthen substrate flooring are optimal (Evink 2002)...... For small mammals, pipe culverts from 0.3m—1 m in diameter are preferable (Clevenger et al. 2001; McDonald & St Clair 2004).................In other words, "one size does not fit all species".................LOS ANGELES OFFICIALS: PLEASE NOTE THIS AS WE GET CLOSER TO THE REALITY OF CREATING A FREEWAY 101/CHESBRO EXIT SAFE HARBOR CROSSING FOR PUMAS, COYOTES, FOXES, BLACK BEARS, BOBCATS, DEER AND THE OTHER MYRIAD SMALLER MAMMALS THAT CALL GREATER LOS ANGELES HOME

http://www.malibutimes.com/news/article_461ae1a2-4419-11e4-ad83-8bea3c795f85.html#.VCOFCUekcqM.email

Wildlife Advocates Launch Campaign to Save Cougars

By expert estimates, the Santa Monica Mountains have less than 10 mountain lions. Advocates hope a $10-million nature overpass could help the population.
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More than 300 people rallied together near Liberty Canyon in Agoura Hills on Friday with a mission — and a hashtag — to protect Southern California’s mountain lions.

“We need to speak for wildlife because they can’t speak for themselves,” said Collin O’Mara, president of the National Wildlife Federation. “We’re creating a model that can be replicated in other parts of the country... This isn’t just an LA story. This is very much a national story.”


State, city and local representatives joined with concerned citizens, activists and fourth-grade students from Woodlake Elementary to show support for a new campaign deemed “#saveLAcougars.” The cause advocates for building a $10-million wildlife crossing that could connect animal habitats divided from freeway developments and population growth, enabling genetic diversity for a dwindling mountain lion population. 
The wildlife overpass is still waiting approval from Caltrans and privately funded grants. While fencing and revegetation projects could begin as early as this year, a wildlife overpass won’t be likely until 2018.
In addition to helping preserve the species, advocates hope the nature overpass could help other animals access green areas needed to survive as well as serve as a community path for bikers, hikers and equestrians.
The Santa Monica Mountains contain less than 10 mountain lions, according to expert estimates, which has led to extremely low genetic diversity and inbreeding.






Skeptics of the project have doubted whether or not an animal would know to use the crossing or what kind of crossing to even build.
“It’s all based on years of research,” said Kate Kuykendall, public affairs officer for National Park Service (NPS). “I don’t think anybody can make any guarantees for sure. We’re going to put the best science behind determining what kind of crossing, where it should go, what kind of cues to give animals.” 
Environmentalists believe the rural landscape and smaller populations near Malibu Canyon and Agoura roads serves as the easiest access point for animals to cross the freeway into new territories. Plans for small modifications and clean-up projects near the freeways may further help the animals cross streets without harming animals or people




.
“The main thing that’s occurring is, this whole fencing is coming out and we’re going to revegetate. When the animals get across, they’ll get back toward the natural area to the left,” Seth Riley, an expert with NPS, said. 
Projects to move fencing and revegetate the area near the Liberty Canyon exits on the U.S. 101 Highway could begin within the next few months, according to Riley. Soon after, activists hope plans for the wildlife crossing will go into effect.
The wildlife crossing seems to have gained traction with many local elected officials, who showed their support on Friday.




State Sen. Fran Pavley noted that 30 years ago, developers had slated the land to be 400 apartment buildings and a carpet store.
“There’s been a lot of good work done behind the scenes. We’ve made some critical planning decisions in this area to help in making sure that this becomes a wildlife corridor for the wildlife and enjoyment of others,” Pavley said. “This could be the longest wildlife corridor in the nation with the help of Caltrans.” 
Malibu City Councilman Lou La Monte stood with the children from Woodlake Elementary when he said, “A wildlife crossing in the most populous county in the United States would send a message to the world that wild things are welcome everywhere.”




Linda Parks, chair of the Santa Monica Mountains Conservancy (SMMC), said the groups involved have already received a $250,000 grant toward the wildlife overpass.
Barbara Marques, Senior Environmental Planner with Caltrans has been working alongside NPS and SMMC for about 10 years now, studying ways to better accommodate the changing climate of development.
In addition to implementing fencing projects on other freeways, a local tunnel project has seen positive wildlife responses and Marques hopes Caltrans can mirror that with the Liberty Canyon crossing.





“We’re going to be implementing some fencing here immediately in the next few months to enhance wildlife connectivity. We’re looking forward to working with our partner agencies on this important project. Maybe the next time we all get together, we’ll be at the groundbreaking for this project.”
---------------------------------------------------------------------------------------------------------------------------

 wildlife corridor important facts


Attention Mary Wiesbrook and colleauges:........note that research to date suggests that mountain lions utilize wildlife underpasses, not overpasses...........let us keep this in mind as we collect the ten million dollars needed to build our freeway 101 wildlife mitigation corridor......see below in the paragraph titled OVERPASSES



Removing and mitigating barriers to wildlife movement

Although roads and urban areas usually occupy only a small fraction of a linkage design, their impacts threaten to block animal movement between the habitat blocks. In this section, we review the potential impacts of these features on ecological processes, identify specific barriers in the linkage design, and suggest appropriate mitigations.
While roads and fences impede animal movement, and the crossing structures we recommend are important, we remind the reader that crossing structures are only part of the overall linkage design. To restore and maintain connectivity between any two wildland blocks, it is essential to consider the entire linkage design, including conserving the land in the linkage. Indeed, investment in a crossing structure would be futile if habitat between the crossing structure and either wildland block is lost.

Impacts of roads on wildlife

While the physical footprint of the nearly 4 million miles of roads in the United States is relatively small, the ecological footprint of the road network extends much farther. Direct effects of roads include road mortality, habitat fragmentation and loss, and reduced connectivity, and the severity of these effects depends on the ecological characteristics of a given species. Direct roadkill affects most species, with severe documented impacts on wide-ranging predators such as the cougar in southern California, the Florida panther, the ocelot, the wolf, and the Iberian lynx (Forman et al. 2003).
In a 4-year study of 15,000 km of road observations in Organ Pipe Cactus National Monument, Rosen and Lowe (1994) found an average of at least 22.5 snakes per km per year killed due to vehicle collisions. Although we may not often think of roads as causing habitat loss, a single freeway (typical width = 50 m, including median and shoulder) crossing diagonally across a 1-mile section of land results in the loss of 4.4% of habitat area for any species that cannot live in the right-of-way. Roads cause habitat fragmentation because they break large habitat areas into small, isolated habit patches which support few individuals; these small populations lose genetic diversity and are at risk of local extinction.
In addition to these obvious effects, roads create noise and vibration that interfere with ability of reptiles, birds, and mammals to communicate, detect prey, or avoid predators. Roads also increase the spread of exotic plants, promote erosion, create barriers to fish, and pollute water sources with roadway chemicals (Forman et al. 2003). Highway lighting also has important impacts on animals (Rich and Longcore 2006).

Mitigation for roads

Wildlife crossing structures that have been used in North America and Europe to facilitate movement through landscapes fragmented by roads include wildlife overpasses & green bridges, bridges, culverts, and pipes. While many of these structures were not originally constructed with ecological connectivity in mind, many species benefit from them (Clevenger et al. 2001; Forman et al. 2003). No single crossing structure will allow all species to cross a road. For example rodents prefer to use pipes and small culverts, while bighorn prefer vegetated overpasses or open terrain below high bridges. A concrete box culvert may be readily accepted by a mountain lion or bear, but not by a deer or bighorn sheep. Small mammals, such as deer mice and voles, prefer small culverts to wildlife overpasses (McDonald & St Clair 2004).

OVERPASSES

Wildlife overpasses are most often designed to improve opportunities for large mammals to cross busy highways. Approximately 50 overpasses have been built in the world, with only 6 of these occurring in North America (Forman et al. 2003). Overpasses are typically 30 to 50 m wide, but can be as large as 200 m wide. In Banff National Park, Alberta, grizzly bears, wolves, and all ungulates (including bighorn sheep, deer, elk, and moose) prefer overpasses to underpasses, while species such as mountain lions prefer underpasses (Clevenger & Waltho 2005).

UNDERPASSES

Wildlife underpasses include viaducts, bridges, culverts, and pipes, and are often designed to ensure adequate drainage beneath highways. For ungulates such as deer that prefer open crossing structures, tall, wide bridges are best. Mule deer in southern California only used underpasses below large spanning bridges (Ng et al. 2004), and the average size of underpasses used by white-tailed deer in Pennsylvania was 15 ft wide by 8 ft high (Brudin 2003). Because most small mammals, amphibians, reptiles, and insects need vegetative cover for security, bridged undercrossings should extend to uplands beyond the scour zone of the stream, and should be high enough to allow enough light for vegetation to grow underneath. In the Netherlands, rows of stumps or branches under crossing structures have increased connectivity for smaller species crossing bridges on floodplains (Forman et al. 2003).

CULVERTS

Drainage culverts can mitigate the effects of busy roads for small and medium sized mammals (Clevenger et al. 2001; McDonald & St Clair 2004). Culverts and concrete box structures are used by many species, including mice, shrews, foxes, rabbits, armadillos, river otters, opossums, raccoons, ground squirrels, skunks, coyotes, bobcats, mountain lions, black bear, great blue heron, long-tailed weasel, amphibians, lizards, snakes, and southern leopard frogs (Yanes et al. 1995; Brudin III 2003; Dodd et al. 2004; Ng et al. 2004). Black bear and mountain lion prefer less-open structures (Clevenger & Waltho 2005). In south Texas, bobcats most often used 1.85 m x 1.85 m box culverts to cross highways, preferred structures near suitable scrub habitat, and sometimes used culverts to rest and avoid high temperatures (Cain et al. 2003). Culvert usage can be enhanced by providing a natural substrate bottom, and in locations where the floor of a culvert is persistently covered with water, a concrete ledge established above water level can provide terrestrial species with a dry path through the structure (Cain et al. 2003). It is important for the lower end of the culvert to be flush with the surrounding terrain. Many culverts are built with a concrete pour-off of 8-12 inches, and others develop a pour-off lip due to scouring action of water. A sheer pour-off of several inches makes it unlikely that many small mammals, snakes, and amphibians will find or use the culvert.
Characteristics which make species vulnerable to the three major direct effects of roads (from Forman et al. 2003).
Effect of roads
Characteristics making a species vulnerable to road effectsRoad mortalityHabitat lossReduced connectivity
Attraction to road habitatx
High intrinsic mobilityx
Habitat generalistx
Multiple resource needsxx
Large area requirements/low densityxxx
Low reproductive ratexxx
Behavioral avoidance of roadsx

ROAD MITIGATION RECOMMENDATIONS

Based on the small but increasing number of scientific studies on wildlife use of highway crossing structures, we offer these standards and guidelines for all existing and future crossing structures intended to facilitate wildlife passage.
  1. Multiple crossing structures should be constructed at a crossing point to provide connectivity for all species likely to use a given area(Little 2003). Different species prefer different types of structures (Clevenger et al. 2001; McDonald & St Clair 2004; Clevenger & Waltho 2005; Mata et al. 2005). For deer or other ungulates, an open structure such as a bridge is crucial. For medium-sized mammals, black bear, and mountain lions, large box culverts with natural earthen substrate flooring are optimal (Evink 2002). For small mammals, pipe culverts from 0.3m—1 m in diameter are preferable (Clevenger et al. 2001; McDonald & St Clair 2004).















  1. At least one crossing structure should be located within an individual's home range. Because most reptiles, small mammals, and amphibians have small home ranges, metal or cement box culverts should be installed at intervals of 150-300 m (Clevenger et al. 2001). For ungulates (deer, pronghorn, bighorn) and large carnivores, larger crossing structures such as bridges, viaducts, or overpasses should be located no more than 1.5 km (0.94 miles) apart (Mata et al. 2005; Clevenger and Wierzchowski 2006). Inadequate size and insufficient number of crossings are two primary causes of poor use by wildlife (Ruediger 2001
















  1. Suitable habitat for species should occur on both sides of the crossing structure (Ruediger 2001; Barnum 2003; Cain et al. 2003; Ng et al. 2004). This applies to both local and landscape scales. On a local scale, vegetative cover should be present near entrances to give animals security, and reduce negative effects such as lighting and noise associated with the road (Clevenger et al. 2001; McDonald & St Clair 2004). A lack of suitable habitat adjacent to culverts originally built for hydrologic function may prevent their use as potential wildlife crossing structures (Cain et al. 2003). On the landscape scale, "Crossing structures will only be as effective as the land and resource management strategies around them" (Clevenger et al. 2005). Suitable habitat must be present throughout the linkage for animals to use a crossing structure




















  1. Whenever possible, suitable habitat should occur within the crossing structure. This can best be achieved by having a bridge high enough to allow enough light for vegetation to grow under the bridge, and by making sure that the bridge spans upland habitat that is not regularly scoured by floods. Where this is not possible, rows of stumps or branches under large span bridges can provide cover for smaller animals such as reptiles, amphibians, rodents, and invertebrates; regular visits are needed to replace artificial cover removed by flood. Within culverts, earthen floors are preferred by mammals and reptiles.

















  1. Structures should be monitored for, and cleared of, obstructions such as detritus or silt blockages that impede movement. Small mammals, carnivores, and reptiles avoid crossing structures with significant detritus blockages (Yanes et al. 1995; Cain et al. 2003; Dodd et al. 2004). In the southwest, over half of box culverts less than 8 x 8 ft have large accumulations of branches, Russian thistle, sand, or garbage that impede animal movement (Beier, personal observation). Bridged undercrossings rarely have similar problems.

















  1. Fencing should never block entrances to crossing structures, and instead should direct animals towards crossing structures (Yanes et al. 1995). In Florida, construction of a barrier wall to guide animals into a culvert system resulted in 93.5% reduction in roadkill, and also increased the total number of species using the culvert from 28 to 42 (Dodd et al. 2004). Fences, guard rails, and embankments at least 2 m high discourage animals from crossing roads (Barnum 2003; Cain et al. 2003; Malo et al. 2004). One-way ramps on roadside fencing can allow an animal to escape if it is trapped on a road (Forman et al. 2003)

















  1. Raised sections of road discourage animals from crossing roads, and should be used when possible to encourage animals to use crossing structures. Clevenger et al. (2003) found that vertebrates were 93% less susceptible to road-kills on sections of road raised on embankments, compared to road segments at the natural grade of the surrounding terrain.
Manage human activity near each crossing structure. Clevenger &Waltho (2000) suggest that human use of crossing structures should be restricted and foot trails relocated away from structures intended for wildlife movement. However, a large crossing structure (viaduct or long, high bridge) should be able to accommodate both recreational and wildlife use. Furthermore, if recreational users are educated to maintain utility of the structure for wildlife, they can be allies in conserving wildlife corridors. At a minimum, nighttime human use of crossing structures should be restricted.

Wednesday, September 24, 2014

Janet Kessler's COYOTE YIPPS Blog is one informative "SONGDOG" site and I recommend all of you checking it out often.............Janet gave me the green light to share her thoughts on the keen vision and intellect of the Coyote--Enjoy her column





From: Coyote Yipps@wordpress.com

Subject: [New post] Coyote Night Vision and What Coyotes Know About Human Vision


WordPress.com
yipps posted: " Coyotes may be seen at any time of the day -- they are diurnal animals, however for convenience in urban areas they have arranged their daily schedules to avoid human activity, so in cities they are active mostly at night when we are not. They can do "


Tuesday, September 23, 2014

The Politics surrounding Wolf Management could be turned into a daytime soap opera................First the Feds control everywhere in America,,,,,,,,,,,Then delisting occurs in parts of the Northern Rockies,,,,,,,,,,,,,,,,,,,,Then there is all types of back and forth as Listed or delisted with delisting up until now the Northern Rocky paradigm(except in the Northwest where some counties are delisted, some are not)...............Then voters in Michigan stop the 2014 hunt until a vote is taken to determine future killing plan(or not hopefully),,,,,,,,,,,And over the last month, the Red Wolf Recovery Program in North Carolina is being re-evaluated and could be dismantled..............And today, Wyoming is stripped of its management of Wolves until they at least provide a Montana/Idaho killing paradigm that allows 15 breeding pairs and 150 animals alive annually.............Note that this capricious "15 pairs/150 wolf paradigm does not have "anchor" as a scientific long term persistent recovery number,,,,,,,,,,,,,,,,Instead, this is what the Feds thought they could get accomplished when Wolves were first being rewilded to the Greater Yellowstone and Idaho region back in the mid 1990's...............A lousy compromise forged with ranching and hunting interests, with the Wolves "not given a vote" in this determination.....................So now we will see how long it will take for Wyoming to agree to the 15 breeding pair/150 wolf number,,,,,,,,,,,,,,,,,,,,,,,,,All this going on while the USFW folks are considering stripping all wolves across the USA of Protection,,,,,,,,,,,,,even though there is viable Wolf habitat across New England, the Appalachian spine and the south which is yet to be repopulated with Wolves(whether they be Gray/Red/Mexican)..........................Whether it be Wolf restoration or international geopolitics,climate change mitigation, job creation, education excellence, health care, immigration et al.................We seem so out of control and unable to forge timely and appropriate rules and actions that reflect our standing as the Leader of the World...................LETS GO USA,,,,, WE ROOT FOR YOU TO ONCE AGAIN BE THE DECISIVE QUARTERBACK AND FRANCHISE # 4 HITTER IN THE LINEUP, reaching informed, educated and "feel right" decisions that the rest of the world will embrace to optimize life here on Planet Earth

https://www.google.com/url?rct=j&sa=t&url=http://www.spokesman.com/blogs/outdoors/2014/sep/23/court-reinstates-endangered-status-wyoming-wolves/&ct=ga&cd=CAIyGjNiY2FkZTRjYzk3Y2I0MjI6Y29tOmVuOlVT&usg=AFQjCNEj7vR1eVmOxV2CvtzdUSe1SZb1qA



Court reinstates endangered status for Wyoming wolves

Although wolf hunting is legal in Wyoming, these hunters go further to make a statement about their attitude toward wolves and their impact on big-game populations in October 2013. (Wind River Outdoor World)
Although wolf hunting is legal in Wyoming, these hunters go further to
 make a statement about their attitude toward wolves and their impact
 on big-game populations in October 2013. (Wind River Outdoor World)

ENDANGERED SPECIES — Federal protections for
 gray wolves in Wyoming were reinstated today after a judge invalidated the U.S. Fish and Wildlife Service’s
 2012 statewide Endangered Species Act delisting of
the species, according to Earthjustice.
The ruling from the U.S. District Court halts the
management of wolves by Wyoming, a state with
a history of anti-wolf policies.

Monday, September 22, 2014

A landmark settlement in 2011 between the Center For Biological Diversity and the Fish and Wildlife Service set legally binding deadlines for the Service to make protection decisions for 757 species around the country. ...........To date, 137 species have gained Endangered Species Act protection as the result of the agreement, and another six have been proposed for protection...........The agreement also allows the Center to seek expedited protection decisions for 10 additional species per year when the agency has missed legal deadlines................ The Alexander Archipelago wolf was one of this year’s 10......................Heavily reliant on old-growth forests in Alaska, , Alexander Archipelago wolves den in the root systems of very large trees and hunt mostly Sitka black-tailed deer, which are themselves dependent on high-quality, old-growth forests, especially for winter survival................. A long history of clearcut logging on the Tongass and private and state-owned lands has devastated much of the wolves’ habitat on the islands of southeast Alaska............. The Forest Service continues to hold timber sales in important wolf habitat..................Let us see if the Feds show the proper empathy for these Wolves and get them the protection they need to persist into the forever

View the page here: http://www.biologicaldiversity.org/news/press_releases/2014/alexander-archipelago-wolf-09-22-2014.html

The Alexander Archipelago Wolf: A Conservation Assessment

The Alexander Archipelago wolf is a subspecies of the gray or timber wolf and is found in southeast Alaska. The information that follows is from the referenced report by Person et al. (1996). This technical report was published by U.S. Department of Agriculture, Forest Service.

Person, David K.; Kirchhoff, Matthew; Van Ballenberghe, Victor; Iverson, George C.; Grossman, Edward. 1996. The Alexander Archipelago wolf: a conservation assessment. Gen. Tech. Rep. PNW-GTR-384. Portland, OR: U.S. Department of Agriculture, Forest Service, Pacific Northwest Research Station. 42 p. (Shaw, Charles G., III, tech. coord.; Conservation and resource assessments for the Tongass land management plan revision).

ABSTRACT

Canis lupus ligoni We summarized the scientific information available for the Alexander Archipelago wolf (Canis lupus ligoni) in the Tongass National Forest of southeast Alaska. Information concerning the morphology, distribution, taxonomy, genetics, and ecology of wolves is presented. Three issues for the conservation of wolves in southeast Alaska are discussed: loss of long-term carrying capacity for deer due primarily to extensive timber harvesting, increased mortality of wolves associated with improved human access from roads, and continued high levels of harvest of wolves by humans. Continued timber harvesting at current levels and by current methods will likely have adverse consequences for some segments of the wolf population. Although some short-term regulatory changes and the management of road access may need to be considered to keep wolf harvest at a sustainable level, the most important consideration is to maintain long-term carrying capacity for deer, the principal prey for most of the wolf population. A series of old-growth forest reserves may provide an effective strategy to increase the likelihood that wolves will persist where extensive timber harvesting has occurred, or is planned.


Keywords: Alexander Archipelago wolf, Canis lupus ligoni, effects of logging on wildlife, population dynamics of wolves, predator-prey dynamics, roads and wolf mortality, Tongass National Forest, southeast Alaska.

SUMMARY

The Alexander Archipelago wolf (Canis lupus ligoni) occupies most of southeast Alaska from Yakutat Bay to Dixon Entrance except for Admiralty, Baranof, and Chichagof Islands. Based on common cranial characteristics, the Alexander Archipelago wolf was considered by early taxonomists to be a distinct subspecies. Recent taxonomic work suggests that these wolves may have originated from a larger subspecific group (C. l. nubilus) that at one time inhabited most of the contiguous Western United States. Wolves probably entered southeast Alaska sometime after the Wisconsin glaciation, following the northward expansion of black-tailed deer (Odocoileus hemionus) along the coast. The hypothesis of a southern origin is supported by recent genetic research showing that wolves in southeast Alaska share a common allele not found in a sample of wolves from interior Alaska or the Yukon. The population is relatively isolated from other wolf populations by water and mountain barriers.

A study of the ecology of wolves in southeast Alaska was conducted on Prince of Wales and Kosciusko Islands from 1992 to 1995. Average home-range size of radio-telemetered wolves was 280 square kilometers (109 mi²), with 75 percent of the radio locations for each pack within “core areas” averaging 124 square kilometers (48 mi²). Pack sizes ranged from 2 to 12, with 7 to 9 typical in early autumn. Annual rates of dispersal averaged 39 percent; 71 percent of dispersers were adults 2 years old and older. Dispersal distances were relatively short (13 to 182 kilometers [5 to 71 mi]) presumably due to inter-island water barriers. Wolf movements were more restricted during the denning and pup-rearing season (mid-April through August), when home ranges were 50 percent smaller than in winter. Of the 14 dens located in this study, all were in old-growth forest within 100 meters (328 feet) of fresh water. One den was under a large log; all others were in cavities beneath the roots of large trees.




Sitka black-tailed deer (O. h. sitkensis) were the primary prey of wolves. Deer remains occurred in 90 percent of wolf feces (scats) examined from Prince of Wales Island. Deer occurred exclusively in 45 percent of the scats. The only other prey occurring with >10 percent frequency was beaver (Castor canadensis). Other prey consumed in small quantities included black bears (Ursus americanus), mustelids, other small mammals, birds, and salmon (Oncorhynchus spp.). Using information on diet composition, consumption rates, and body size of prey, we estimated that wolves on islands in southern southeast Alaska consumed an average of 26 deer per wolf per year (SE = 4.1).

Most of the wolves in southeast Alaska occur on the large islands south of Frederick Sound. These islands (game management units 2 and 3) support approximately 60 to 70 percent of the total population. By extrapolating from empirical population estimates for Prince of Wales Island, we estimated the autumn 1994 population of wolves in southeast Alaska at slightly over 900 animals (SE = 216). Hunting, trapping, and illegal killing accounts for a high percentage of the mortality in wolves. Based on analysis of trapping and hunting morality by wildlife analysis area (WAA), we determined that mortality was correlated with the linear kilometers of road within WAA's. Indeed, reported wolf harvest increased twofold when the length of road below 370 meters (1200 ft) elevation exceeded 95 kilometers (59 mi), regardless of size of the WAA. This corresponded to an approximate road density of 0.4 kilometer per square kilometer (0.7 mi/mi²), most of which were open to human access. Between 1993 and 1995, the average annual mortality in a total sample of 24 radio-collared wolves on Prince of Wales Island was 50 percent SE = 13 percent). If applied to the overall wolf population on Prince of Wales Island, this rate of mortality would not be sustainable.



Wolf populations are closely tied to population levels of their ungulate prey. For southeast Alaska, we predicted the number of deer required per wolf to attain equilibrium between deer and wolves by using a Monte Carlo simulation of a model that calculated equilibrium ratios for wolves and their ungulate prey. We assumed a high average finite rate of increase for deer (1.3), a mean predation rate of 26 deer per wolf per year, and a human harvest of deer equal to 21 percent of the annual increment. Our results suggest that 170 to 180 deer per wolf are needed for a 95-percent probability of equilibrium, provided that mortality of deer due to predation is primarily additive. We cannot suggest a minimum deer population because we do not know what constitutes a minimum viable wolf population either demographically or genetically. Nevertheless, if we expect to sustain the current postdenning population of 250-300 wolves on Prince of Wales Island (along with subsistence and sport harvests of deer) with a high probability of attaining equilibrium, then sufficient habitat is needed to support 42,500 to 54,000 deer.






Our review raises a number of issues concerning the long-term sustainability of wolves in southeast Alaska. Many more data are needed on wolf population structure, genetic structure, and predator-prey relations to fully address these issues and the overall question of viability. The Alexander Archipelago wolf exists in small numbers in a rapidly changing insular environment. Projected growth in human population, increasing road access, and the continuing loss and fragmentation of high-quality deer habitat will increase the risk of not maintaining a viable, well-distributed population of wolves in southeast Alaska. The area of most immediate concern is game management unit 2, including Prince of Wales and Kosciusko Islands.

Management actions that address risks to wolf populations include modifying hunting and trapping regulations as necessary, limiting construction of new roads and effectively closing some existing ones, and modifying timber harvest strategies to minimize fragmentation and loss of critical deer winter range. Habitat to support a minimum density of 5 deer per square kilometer (13 deer/mi²), where deer are the primary prey for wolves, would provide for current levels of deer harvest by hunters, trappers, and wolves. In areas less productive for deer, maintaining current densities of deer is particularly important. Setting aside contiguous blocks of habitat within each biogeographic province that are large enough to encompass at least one wolf pack core home range (200 square kilometers [76 mi²]) would markedly increase the likely persistence of wolves, especially if the reserves contain high-quality deer habitat sufficient to support an average density of deer equal to 7 deer per square kilometer (18 deer/mi²). The objective of maintaining undisturbed blocks of habitat within each biogeographic province is to assure the persistence of several wolf packs that will serve as source populations capable of replacing wolves that periodically disappear from adjacent disturbed lands.





PURPOSE AND NEED

In 1990, wolves in southeast Alaska were identified by a USDA Forest Service-sponsored interagency committee as a species for which there may be concerns about viability or distribution as a result of extensive timber harvesting in the Tongass National Forest. In December 1993, the Biodiversity Legal Foundation (Boulder, CO) and an independent biologist from Haines, AK, filed a petition with the U.S. Fish and Wildlife Service (FWS) requesting that wolves in southeast Alaska be listed as a threatened subspecies pursuant to the Endangered Species Act of 1973, as amended. The FWS ruled that listing was not warranted at this time, but added: “However, it is clear by our analysis that without significant changes to the existing Tongass Land Management Plan, the long-term viability of the Alexander Archipelago wolf is seriously imperiled.” Following that notice, this assessment was prepared under the auspices of the memorandum of understanding among the Forest Service, Alaska Department of Fish and Game, and the Fish and Wildlife Service. They, along with the Regional Forester, Alaska Region, requested that a conservation assessment be prepared to summarize available scientific information on the status and ecology of wolves in southeast Alaska, and to identify issues regarding their long-term viability and distribution.






This conservation assessment was chartered under the Tongass [National Forest] Land Management Plan revision and the interagency memorandum of understanding (MOU) among the Alaska Region of the Forest Service, the Alaska Region of the U.S. Fish and Wildlife Service, and the Alaska Department of Fish and Game to conserve species tending toward listing. Our report summarizes the most current information available on the taxonomy, genetics, distribution, ecology, and population dynamics of wolves in southeast Alaska. Much of the information has not been published. In particular, we have drawn heavily from ongoing studies on wolves in southeast Alaska that is cosponsored by the USDA Forest Service, the Alaska Department of Fish and Game, the U.S. Fish and Wildlife Service, and the University of Alaska, Fairbanks. Results from that work are preliminary.